To cite memes in publications use:

  Nystrom, SL, McKay, DJ (2021) Memes: A motif analysis environment in
  R using tools from the MEME Suite.  PLOS Computational Biology 17(9):
  e1008991. https://doi.org/10.1371/journal.pcbi.1008991

A BibTeX entry for LaTeX users is

  @Article{,
    title = {Memes: A motif analysis environment in R using tools from the MEME Suite},
    author = {Spencer L. Nystrom and Daniel J. McKay},
    journal = {PLOS Computational Biology},
    year = {2021},
    doi = {10.1371/journal.pcbi.1008991},
    volume = {17},
    pages = {1-14},
    url = {https://doi.org/10.1371/journal.pcbi.1008991},
  }